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黄土丘陵沟壑区抗侵蚀植物及其群落特征研究
寇 萌
学位类型博士
导师焦菊英
2016-05
学位授予单位中国科学院研究生院
学位授予地点北京
关键词抗侵蚀植物 植被演替 群落结构 生态位宽度 土壤侵蚀
摘要

黄土丘陵沟壑区水土流失严重,土壤侵蚀严重干扰着植物的发育与演替过程。在
土壤侵蚀严重地区生存的植物,采用不同的生存策略适应和抵抗土壤侵蚀造成的各种
胁迫与干扰,并能发展为可抵抗土壤侵蚀的植物群落。本研究基于对黄土丘陵沟壑区
延河流域多年植被调查资料的整理分析,根据抗侵蚀植物定义及 Braun-Blanquet 植物
社会学方法,筛选该区潜在的抗侵蚀植物,探讨其适应机制及其在群落中的作用,分
析不同抗侵蚀植物的演变规律;通过对不同侵蚀环境下植物群落的调查与分析,阐明
植被恢复过程中不同抗侵蚀植物群落改善土壤侵蚀环境的能力;通过计算不同抗侵蚀
植物群落结构化植被因子指数,并结合其土壤侵蚀量数据,探明能够抵抗土壤侵蚀的
群落结构特征。主要研究结果如下:
1)共筛选出潜在抗侵蚀植物 42 种,分属 18 科 33 属。禾本科、豆科、菊科、蔷
薇科在研究区占优势地位,是退耕地植被恢复演替过程中的主要植物组成。该区气候
的自然选择决定了地区植物生长型以灌、草为主,灌木多为高位芽或地上芽植物,多
年生草本植物以地面芽为主;水分生态类型以中生、旱生为主。
2)42 种抗侵蚀植物通过土壤种子库、植冠种子库、种子萌发休眠策略、幼苗建
植及营养繁殖等方式维持自身的生存繁衍;通过解剖形态上的特殊结构、厚的保护组
织、发达的栅栏组织以及生理上的抗氧化机制和渗透调节作用中的一种或是数种机制
来适应侵蚀环境。不同抗侵蚀植物生态位宽度在 0.023-0.359 之间,抗侵蚀植物在适
宜生境中可作为群落优势种或伴生种,在其它生境条件下是群落偶见种。
3)在自然恢复过程中,草本群落演替后期抗侵蚀植物开始出现生境分化;延河
流域南部森林带、中部森林草原带的沟坡与沟沿线等局部区域分别有乔、灌木的分布,
北部草原带则少见灌木。在刺槐林下植被恢复过程中,林下出现的抗侵蚀植物较少,
只有部分一年生杂草和禾本科植物的存在;人工林逐渐衰退后,演替中后期的植物开
始成为林下植被的优势种。在人工灌木林生长相对稳定的时期,林下抗侵蚀植物并没
有出现更替。
4)由较多物种形成的群落结构尤为重要,不同物种组成的地上和地下复杂层次
结构才能起到有效改善土壤侵蚀环境的作用。抗侵蚀植物群落的改善能力与群落所处的生境密切相关,在延河流域南部森林带,乔、灌、草形成的群落层次结构及较多的
枯落物是改善土壤环境的主要因素;在中部森林草原带和北部草原带,植物细根的作
用是改善土壤环境的关键因素。
5)不同群落类型有各自能有效减少土壤侵蚀强度的结构化植被因子指数 Cs。演
替早期群落及人工林未表现出较强的减蚀能力,当 Cs 分别大于 20%和 30%左右,且
草本层和枯落层盖度较大时,土壤侵蚀强度小于 2500 t km -2 yr -1 。演替中、后期群落
及自然乔、灌群落 Cs 分别大于 30%和 36%,且近地面的灌木、草本层及贴地面的枯
落层中至少有一层盖度较大时,可有效减少土壤侵蚀,尤其在暴雨年份,禾本科植物
群落及自然乔、灌群落能有效控制土壤侵蚀,当 Cs 分别大于 38%和 30-40%,且近地
面至少有一层盖度较大时,土壤侵蚀强度小于黄土高原地区允许土壤流失量。不同水
文年群落能有效抵抗土壤侵蚀的 Cs 值有明显区别,暴雨年份植物群落有效减少土壤
侵蚀所需的 Cs 值较高。
关键词 :抗侵蚀植物;植被演替;群落结构;生态位宽度;土壤侵蚀

其他摘要

The Loess Plateau is well known for its severe soil erosion. Soil erosion severely
interferes with the process of plant development and succession. In this serious soil erosion
region, plants can overcome stress and disturbance due to soil erosion through different
survival strategies, and developed into plant communities that could resist soil erosion.
This study was based on several years’ vegetation survey data of the Yan River basin in the
hilly and gully region of the Loess Plateau. The definition of erosion-resistant species and
Braun-Blanquet phytosociology methods were used to select potential erosion-resistant
species. The community species composition of different recovery stages, plant growth and
reproductive form characteristic, and plant niche breadth were used to explore its
adaptation mechanism and its role in the plant community, and to analyze the evolvement
rule of different erosion-resistant species. Plant community survey and analysis data under
different soil erosion environments were used to elucidate the ability of different
erosion-resistant plant communities to improve soil erosion environment in the vegetation
recovery process. Plant community structure vegetation cover index and soil erosion data
in different erosion-resistant plant communities were used to illuminate the community
structure characteristics that could resist soil erosion. The main results are as follows:
1) A total of 42 potential erosion-resistant species were selected, which belong to 18
families and 33 genera. Gramineae, Leguminosae, Compositae, Rosaceae occupied the
dominant position in the study area; they were the main species in vegetation restoration
succession process. Due to the climate in this region, the plant growth form was given
priority to with shrub and herbs. Shrubs were mostly phanerophyte and chamaephytes, and
herbs were dominant by hemicryptophytes. Plant water ecological type was given priority  to with mesophyte and xerophytes.
2) Forty-two potential erosion-resistant species maintained their own survival and
reproduction through soil seed bank, plant crown seed bank, seed germination and
dormancy strategies, seedling planting and vegetative propagation. They could adapt to
environmental erosion by one or several mechanism of the special structure of anatomical
morphology, thick protective tissue, developed palisade tissue, antioxidant mechanism and
osmotic adjustment. The ecological niche breadth of different erosion-resistant species was
from 0.023 to 0.359. The erosion-resistant species could be regarded as dominant species
or accompanying species in the suitable habitat, it was an occasional species in other
habitats.
3) In the process of natural vegetation recovery, the erosion-resistant species began to
habitat differentiation in herbaceous community late succession stages. Tree and shrub
distributed in local region along gully slope, shoulder line of valley, etc of forest zone and
forest steppe zone of Yan river basin, respectively. Shrub rarely appeared in steppe zone of
Yan river basin. In the process of understory vegetation restoration under Robinia
pseudoacacia plantation, the erosion-resistant species was less, only some annual weeds
and Gramineous plants appeared under plantation. After Robinia pseudoacacia plantation
decline, the mid and late stage species were the dominant species under plantation. During
the period of relatively stable growth of the shrub plantation, there was no replacement of
erosion-resistant species under plantation.
4) The suitable community structure, which was formed by many species, was
particularly important. The ground and underground complex hierarchical structure formed
by different species could effectively improve soil environment. In addition, the
improvement ability of erosion-resistant plant community was closely related to its habitat.
In forest zone, phytocenosium structure formed by tree, shrub, herbs, and more litter were
the main factors to improve soil environment. In forest steppe zone and steppe zone, plant
fine root was the key factor to improve soil environment.
5) Different communities have their own suitable structure vegetation cover index (Cs)
that could effectively reduce soil erosion intensity. Early succession community and
afforestation did not show a strong erosion-resistant, when the Cs were more than 20% and
30% respectively, and that the cover of herb layer and litter layer was higher, the soil  erosion intensity was less than 2500 t km -2 yr -1 . When Cs of mid and late succession
communities, and natural tree and shrub communities were more than 30%, and 36%
respectively, and that at least one of the cover of near surface layers (shrub, herb, litter
layer) was higher, these communities could be effective in reducing soil erosion. Especially
in the year of rainstorm, Gramineous plant communities and natural tree and shrub
communities could effectively control soil erosion, when the Cs of these communities were
more than 38% and 30-40%, respectively, and that at least one of the cover of near surface
layers was higher, soil erosion intensity was smaller than soil loss tolerance in the Loess
Plateau Region. The Cs value that community could effectively resist soil erosion had
obvious difference in different hydrological years. In the year of rainstorm, the
communities need to have higher Cs value when it was able to effectively reduce soil
erosion.
Key Words: erosion-resistant plant; vegetation succession; community structure
characteristics; ecological niche breadth; soil erosion

语种中文
文献类型学位论文
条目标识符sbir.nwafu.edu.cn/handle/361005/9099
专题水保所知识产出(1956---)
推荐引用方式
GB/T 7714
寇 萌. 黄土丘陵沟壑区抗侵蚀植物及其群落特征研究[D]. 北京. 中国科学院研究生院,2016.
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